S húa bi trung tõm hc liu Header Page of 126 http://www.lrc.tnu.edu.vn/ I HC THI NGUYấN TRNG I HC NễNG LM LNG VN HO NGHIấN CU THNH PHN LOI V C IM PHN B CA THN MM CHN BNG (Gastropoda) TRấN CN RNG THCH THNH (THANH HO) THUC VN QUC GIA CC PHNG Chuyờn ngnh: Lõm hc Mó s: 60 62 02 01 LUN VN THC S KHOA HC LM NGHIP Ngi hng dn khoa hc: PGS.TS VN NHNG Thỏi Nguyờn, 2013 Footer Page of 126 Header Page of 126 i LI CAM OAN Tụi xin cam oan bn lun ny l cụng trỡnh nghiờn cu thc s ca cỏ nhõn, c thc hin trờn c s nghiờn cu thc a rng Thch Lõm, Thch Thnh (Thanh Húa) thuc Vn Quc gia Cỳc Phng, Vit Nam di s hng dn khoa hc ca PGS TS Vn Nhng Cỏc s liu v kt qu ca lun l trung thc, khỏch quan v cha c cụng b bt c cụng trỡnh nghiờn cu no khỏc Thỏi Nguyờn, thỏng 10 nm 2013 Tỏc gi Lng Vn Ho Footer Page 2S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 ii LI CM N hon thnh lun trc tiờn tụi gi li cm n sõu sc ti thy giỏo PGS TS Vn Nhng ó trc tip hng dn, truyn t kin thc thc t, phng phỏp lun, ụn c kim tra sut quỏ trỡnh nghiờn cu Tụi cng xin gi li cm n ti Ban giỏm c Vn Quc gia Hong Liờn, c bit ụng Phm Vn ng, Giỏm c Vn ó to iu kin thun li tụi c tham gia hc tp; cm n ụng Trng Quang Bớch v ụng Vn Lp, Ban giỏm c VQG Cỳc Phng ó to iu kin, giỳp quỏ trỡnh nghiờn cu thc a, cung cp cỏc ti liu tham kho vit lun Ngoi ra, tụi cng vụ cựng bit n cỏc em sinh viờn, Lu Th Thanh Hng (i hc S phm H Ni) ó giỳp rt nhiu phõn tớch mu, Lng Vn Hong (i hc Nụng Lõm Bc Giang), c Sỏng (nghiờn cu sinh, Ging viờn i hc Tõy Bc) cựng ng nghip v bn bố Hong Trng Giang, Ló Th Qunh, Chu Vn Tn, Nguyn Kim Hong, ngi v v l ng nghip Nguyn Th Hng Thm ó tr giỳp v khớch l tụi quỏ trỡnh thu thp mu vt, ti liu, chnh sa nh, tớnh toỏn s liu phc v vit lun Nhiu s giỳp quý bỏu ca cỏ nhõn, ngi dõn a phng ó tham gia trc tip vo cụng vic thc a, tr li phng vn, to iu kin ni n, ch , qua õy cho tụi gi li cm n chõn thnh ti tt c s giỳp quý bỏu ú Li cm n chõn tỡnh ti cỏc ng nghip nc ngoi, Giỏo s Fred Naggs (Bo tng Lch s T nhiờn London, Anh), Tin s Miklos Szekeres (Vin sinh hc thc vt, Vin hn lõm khoa hc Hungari) ó tr giỳp v kinh phớ thu thp s liu hin trng, cung cp ti liu v giỳp kim tra mu vt, nh tờn mt s loi c cn Thch Lõm, Thch Thnh, Thanh Húa Thỏi Nguyờn, thỏng 10 nm 2013 Tỏc gi Lng Vn Ho Footer Page 3S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 iii MC LC LI CAM OAN LI CM N MC LC DANH MC CC T VIT TT DANH MC CC BNG BIU DANH MC CC HèNH V M U 1 Lý chn ti Mc tiờu ca ti Ni dung v phm vi nghiờn cu í ngha nghiờn cu ca ti Chng 1: TNG QUAN NGHIấN CU 1.1 Tng quan nghiờn cu 1.1 Nghiờn cu Thõn mm Chõn bng trờn th gii 1.1.2 Nghiờn cu Thõn mm Chõn bng Vit Nam 1.1.3 Nghiờn cu VQG Cỳc Phng 1.2 Mt s nột khỏi quỏt khu vc nghiờn cu 10 1.2.1 V trớ a lý 10 1.2.2 c im a hỡnh 10 1.2.3 Khớ hu, thy 12 1.2.3.1 Ch nhit 12 1.2.3.2 Ch ma 12 1.2.3.3 m khụng khớ 12 1.2.3.4 Thy 13 1.2.3.5 c im khu h ng, thc vt Thch Lõm, Thch Thnh 14 1.2.4 Tỡnh hỡnh kinh t v xó hi 16 1.2.4.1 C cu kinh t 16 1.2.4.2 C s h tng 17 1.2.4.3 Tỡnh hỡnh y t, giỏo dc, húa 17 1.2.4.4 Thnh phn dõn tc 17 Chng 2: NI DUNG V PHNG PHP NGHIấN CU 19 2.1 i tng v phm vi nghiờn cu 19 2.1.1 i tng nghiờn cu 19 2.1.2 Phm vi nghiờn cu Thõn mm Chõn bng trờn cn 19 2.2 Thi gian v a im nghiờn cu 20 Footer Page 4S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 iv 2.2.1 Thi gian nghiờn cu 20 2.2.2 a im nghiờn cu 20 2.3 Ni dung v phng phỏp nghiờn cu 21 2.3.1 Ni dung nghiờn cu 21 2.3.1 Phng phỏp nghiờn cu 21 2.3.1.1 Phng phỏp nghiờn cu ngoi thc a 21 2.3.1.2 iu tra, phng dõn a phng 23 2.3.1.3 Phng phỏp phõn tớch phũng thớ nghim 23 2.3.1.4 Gii phỏp thc hin ti 25 Chng 3: KT QU NGHIấN CU V THO LUN 27 3.1 Thnh phn loi v a dng thõn mm chõn bng khu vc nghiờn cu 27 3.1.1 Thnh phn loi khu vc nghiờn cu 27 3.1.2 Mc a dng cỏc loi c cn khu vc nghiờn cu 34 3.1.2.1 a dng loi khu vc nghiờn cu 34 3.1.2.2 c im hỡnh thỏi cỏc loi khu vc nghiờn cu 36 3.2 c im phõn b khu vc nghiờn cu 55 3.2.1 Phõn b loi theo sinh cnh 61 3.2.1.1 Rng gi trờn nỳi ỏ vụi (Rgn) 61 3.2.1.2 Rng th sinh trờn nỳi ỏ vụi (Rtsn) 63 3.2.1.3 Rng nghốo trờn nỳi ỏ vụi (Rnn) 66 3.2.1.4 Rng th sinh trờn i t (Tts) 68 3.2.1.5 Tre, na (Tr) 70 3.2.1.6 Rng tỏi sinh sau nng ry (Rtsnr) 72 3.2.1.7 Sinh cnh nhõn tỏc (Nr) 74 3.2.2 Phõn b theo a hỡnh nỳi ỏ vụi v nỳi t 74 3.2.2.1 Thnh phn loi phõn b theo a hỡnh nỳi ỏ vụi 76 3.2.2.2 Thnh phn loi phõn b theo a hỡnh nỳi t 78 3.3 Giỏ tr s dng 80 3.3.1 Nhn bit v thnh phn loi v tỡnh trng c cn Thch Lõm 80 3.3.2 V giỏ tr s dng lm thc phm v thng mi 81 3.3.3 Giỏ tr y dc 83 KT LUN V KIN NGH 84 Kt lun 84 Kin ngh v xut gii phỏp bo tn 87 TI LIU THAM KHO 89 Footer Page 5S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 v DANH MC CC CM T VIT TT VQG : Vn Quc gia Rgn : Rng gi trờn nỳi ỏ vụi Rtsn : Rng th sinh trờn nỳi ỏ vụi Rnn : Rng nghốo trờn nỳi ỏ vụi Rts : Rng th sinh trờn i t Rtsnr : Rng tỏi sinh sau nng ry Tr : Rng tre na Nr : Nng ry Footer Page 6S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 vi DANH MC CC BNG, BIU Bng 1.1: Cỏc ch tiờu khớ hu c bn khu vc VQG Cỳc Phng 13 Bng 1.2: S lng Taxon cỏc ngnh thc vt bc cao Cỳc Phng 15 Bng 2.1: Phõn chia mc phõn b c cn theo sinh cnh 25 Bng 3.1: Danh sỏch cỏc loi c cn Thch Lõm, Thch Thnh 28 Bng 3.2: Tớnh a dng cỏc loi thuc cỏc h c cn Thch Lõm 34 Bng 3.3: a dng c cn mt s vựng nỳi ỏ vụi Min Bc Vit Nam 35 Bng 3.4: V trớ cỏc ụ nghiờn cu Thch Lõm, Thch Thnh 54 Footer Page 7S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 vii DANH MC CC HèNH V Hỡnh 1.1: S khu vc Vn Quc gia Cỳc Phng 11 Hỡnh 2.1: S v trớ nghiờn cu c cn Thch Lõm 20 Hỡnh 2.2: Hỡnh thỏi ngoi v mt phn bờn ca v c cn 23 Hỡnh 3.1: S cu trỳc phõn loi c cn Thch Lõm, Thch Thnh (VQG Cỳc Phng) 33 Hỡnh 3.2 Biu so sỏnh a dng loi c cn gia mt s vựng nỳi ỏ vụi phớa Bc Vit Nam 36 Hỡnh 3.3 Biu phõn b cỏc loi theo sinh cnh 55 Hỡnh 3.4 Biu phõn b cỏc h theo sinh cnh 57 Hỡnh 3.5 Biu phõn b cỏc h theo a hỡnh 58 Hỡnh 3.6 Biu phõn b cỏc loi trờn c hai kiu a hỡnh 59 Hỡnh 3.7 Biu phõn b loi theo sinh cnh rng gi trờn nỳi ỏ vụi .62 Hỡnh 3.8 Biu phõn b loi theo sinh cnh rng th sinh trờn nỳi ỏ vụi 65 Hỡnh 3.9 Biu phõn b loi theo sinh cnh rng nghốo trờn nỳi ỏ vụi 67 Hỡnh 3.10 Biu phõn b loi theo sinh cnh rng th sinh trờn i t 69 Hỡnh 3.11 Biu phõn b loi theo sinh cnh rng tre na 71 Hỡnh 3.12 Biu phõn b loi theo sinh cnh rng th sinh sau nng ry 73 Hỡnh 3.13 Biu phõn b cỏc loi trờn c hai kiu a hỡnh 75 Hỡnh 3.14 Biu phõn b loi trờn a hỡnh nỳi ỏ vụi 77 Hỡnh 3.15 Biu phõn b loi trờn a hỡnh nỳi t 79 Footer Page 8S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page of 126 M U Lý chn ti Lp Thõn mm Chõn bng (Gastropoda) l mt lp thuc Ngnh thõn mm (Mollusca) cú vai trũ c bit quan trng h sinh thỏi v giỏ tr thc tin i vi ngi Trong lp Chõn bng cú phõn lp: phõn lp Mang trc (Prosobranchia), phõn lp Mang sau (Opisthobranchia) v Phõn lp Cú phi (Pulmonata) Trong phõn lp ny, phõn lp Mang sau hon ton bin, phõn lp mang trc t l loi sng nc chim phn ln cũn mt s cn, phõn lp Pulmonata sng trờn cn Tri qua s tin húa hng triu nm ca Thõn mm Chõn bng ó phỏt sinh nhiu loi v cú s lng loi phong phỳ ch ng th hai sau lp Cụn trựng [159, 174] c bit nhúm cn vi cỏc mụi trng sng c trng ó hỡnh thnh nờn a dng cao Rt nhiu loi s chỳng l ngun thc phm quan trng i vi ngi [2, 166] Cỏch õy khong 7.500 nm, nhng húa thch ging Cyclophorus (Cyclophoridae) ng Ngi Xa Vn Quc gia Cỳc Phng l minh chng ngi c i ó s dng lm ngun thc phm [16] Trong h sinh thỏi, c cn l thnh phn khụng th thiu chui v li thc n, c bit vi mt s loi chim, loi thỳ n tht nh Trong chu trỡnh phõn gii vt cht, c cn l nhúm n thc vt bc thp v mựn bó tng thm mc Tuy nhiờn nhiu loi s chỳng l vt ch trung gian, lan truyn gõy bnh cho ngi v ng vt [164, 165] Ngoi ra, mt s loi cú th phỏ hoi mng (c sờn- Achatina fulica) [165] Vit Nam cỏc nghiờn cu v Thõn mm Chõn bng cũn hn ch, nhiu vựng cha cú dn liu Cỏc nghiờn cu t rt sm nhng kộo di nhiu th k, kt qu nghiờn cu cha phn nh y v a dng, c trng v hỡnh thỏi, kớch thc, phõn loi, phõn b, giỏ tr thc tin Footer Page 9S of húa 126.bi trung tõm hc liu http://www.lrc.tnu.edu.vn/ Header Page 10 of 126 Vn Quc gia Cỳc Phng hin ti b chia ct thnh phn c lp bi sụng Bi v ng H Chớ Minh Phn ln din tớch (18.000 ha) nm phớa ụng ca sụng Bi v khong 2.000 rng nm phớa Tõy sụng Bi [8] Phn phớa Tõy bao gm cỏc xó Thch Lõm (chim i b phn din tớch rng trờn nỳi ỏ vụi) v cỏc xó n Ngha, Tõn M thuc tnh Hũa Bỡnh Vỡ c trng khu vc VQG Cỳc Phng hin ti chia ct thnh vựng nh trờn, cho nờn khuụn kh ca lun ny ch dng li mc nghiờn cu v thnh phn loi v c trng phõn b ca Thõn mm Chõn bng cn khu vc phớa Tõy ca sụng Bi thuc Thch Thnh, Thch Lõm (Thanh Húa), VQG Cỳc Phng Vỡ vy, chỳng tụi la chn ti Nghiờn cu thnh phn loi v c im phõn b ca Thõn mm Chõn bng (Gastropoda) trờn cn rng Thch Thnh (Thanh Húa) thuc Vn Quc gia Cỳc Phng Mc tiờu ca ti - Mc tiờu chung: bc u xõy dng c s thụng tin v a dng khu h c cn, xem xột mi quan h ca chỳng vi sinh cnh t nhiờn v mi liờn quan n cng ng dõn c sng gn rng; ng thi l c s khoa hc v thc tin cho chớnh quyn a phng v Vn Quc gia Cỳc Phng, tnh Ninh Bỡnh cỏc gii phỏp qun lý, bo v v phỏt trin a dng sinh hc - Mc tiờu c th: + Xỏc nh thnh phn v a dng cỏc loi Thõn mm Chõn bng trờn cn khu rng Thch Lõm, Thch Thnh (Thanh Húa) thuc Vn Quc gia Cỳc Phng; + Tỡm hiu c im phõn b ca chỳng trờn cỏc sinh cnh v a hỡnh thuc khu vc nghiờn cu v giỏ tr s dng chỳng; + xut cỏc gii phỏp gim thiu tỏc ng bt li ca cng ng dõn c vựng m vo ti nguyờn rng Cỳc Phng, tnh Ninh Bỡnh Sofhúa bi trung tõm hc liu Footer Page 10 126 http://www.lrc.tnu.edu.vn/ Header Page 94 of 126 86 Clausiliidae, Cyclophoridae, Diplommatinidae, Euconulidae, Helicidae, Pupunidae v Trochomorphidae; cú 21 loi cú mt t 0,003 n 0,008 cỏ th/ m2 thuc 10 h, gm: Achatinidae, Ariophantidae, Bradybaenidae, Camaenida, Clausiliidae, Cyclophoridae, Dyakiidae, Enidae, Euconulidae v Subulinidae + Qun th mt s loi c cn ang c dựng lm thc phm (ging Cyclophorus v Camaena) suy gim khong 50% so vi nm trc Loi c sờn ngoi lai Achatina fulica Fộrussac, 1822 (Achatinidae) ch tỡm thy 01 v trờn khu vc nghiờn cu õy l ghi nhn mi vỡ loi ny cú phõn b rt rng, thớch nghi vi nhiu kiu a hỡnh, sinh cnh khỏc 1.3 Giỏ tr thc tin ca Thõn mm Chõn bng trờn cn Thch Lõm, Thch Thnh (Thanh Húa) thuc VQG Cỳc Phng + Cỏc loi c cn bn a, cú kớch thc ln ang c ngi dõn dựng lm thc phm, thuc ging: Cyclophorus (Cyclophoridae) v Camaena (Camaenidae); mt s loi khỏc ụi cng c dựng lm thc phm, cú kớch thc nh hn, gm: Pollicaria gravida (Pupinidae), Tropidauchenia (Clausiliidae), ớt loi thuc ging Oospira, Phaedusa (Clausiliidae) Loi c ngoi lai Achatina fulica khụng dựng lm thc phm + Mt s loi c cn c cho l cú giỏ tr thc phm cao nht, gm: Camaena vanbuensis Smith, 1898, Camaena gabriellae (Dautzenberg et dHamoven., 1887), Cameana dugasti (Morlet, 1892), Cameana dugasti (Morlet, 1892) Cỏc loi ny cú s lng ớt, him gp v ớt c bỏn th trng Cỏc loi ph bin bỏn th trng l cỏc loi thuc ging Cyclophorus (Cyclophoridae) + Cú 01 loi ngi dõn cho l cú th dựng lm thuc l Geotrochatella jourdyi (Helicinidae), cha cỏc bnh nh: co git st cao, tiu ng, viờm amian, viờm hng, quai b, loa lch (lao hch), nht c, tr sang (tr viờm loột), thoỏt giang (sa trc trng), vt thng cụn trựng cn t + V kinh nghim thu bt, chn nuụi c cn: ngi dõn thng i bt vo ma t thỏng n thỏng c cn bũ ngoi kim n, sinh sn Ngi dõn cha tng nuụi c cn v cha thy nuụi Sofhúa bi trung tõm hc liu Footer Page 94 126 http://www.lrc.tnu.edu.vn/ Header Page 95 of 126 87 Kin ngh v xut gii phỏp bo tn 2.1 Kin ngh + Tỡnh trng khai thỏc lõm sn cũn din mt s ni, mc thp nhng xy ti khu vc rng gi trờn nỳi ỏ vụi vỡ th cn tng cng cụng tỏc kim tra, kim soỏt nhm ngn nga tỡnh trng khai thỏc trỏi phộp lõm sn, õy l mt nhng nguyờn nhõn trc tip dn n thay i cu trỳc, tng tỏn rng, nh hng ti tớnh a dng cỏc loi c cn + Vic khai thỏc trỏi phộp mt s loi c cn cha c kim soỏt cht ch l nguyờn nhõn dn n tỡnh trng suy gim qun th v phõn b ca chỳng, vỡ th cn tng cng giỏm sỏt khai thỏc vo sinh sn, c bit i vi cỏc loi ang c s dng lm thc phm v thng phm ti a phng + Cụng tỏc giỏo dc, tuyờn truyn nõng cao nhn thc v bo tn sinh hc ó c tin hnh nhng khụng c thng xuyờn, liờn tc v cha cú chuyờn sõu v cỏc loi ng vt nh, c cn vỡ th nờn xõy dng chng trỡnh tuyờn truyn chuyờn sõu bo tn loi ng vt nh, thõn mm + Nghiờn cu v sinh hc, sinh thỏi ca c cn Cỳc Phng ó v ang c tin hnh nhng cha thc hin vic i vi mt s loi cú giỏ tr kinh t cao, cú s lng qun th thp vỡ th VQG Cỳc Phng cn tip tc nghiờn cu v m rng i vi cỏc loi cú giỏ tr kinh t, cú s lng ớt + Trong nghiờn cu ny cũn nhiu loi cha xỏc nh c tờn vỡ th tip tc thu hỳt cỏc chuyờn gia tham gia nghiờn cu, VQG Cỳc Phng tng cng nhõn lc nghiờn cu, thu thp b sung mu vt, lu tr ti bo tng Cỳc Phng phc v cụng tỏc giỏo dc v nghiờn cu lõu di + phc v cụng tỏc giỏo dc bo tn mụi trng v a dng sinh hc tt hn, VQG Cỳc Phng nờn t chc in n, xut bn cỏc thụng tin v c cn v ph bin rng rói cho ngi dõn a phng vựng m v lừi VQG Cỳc Phng Sofhúa bi trung tõm hc liu Footer Page 95 126 http://www.lrc.tnu.edu.vn/ Header Page 96 of 126 88 2.2 Gii phỏp bo tn loi + V giỏo dc nõng cao nhn thc: n v bo tn cn kt hp vi ngi dõn v chớnh quyn a phng, c s giỏo dc xõy dng v thc hin, thng xuyờn v liờn tc v bo tn thiờn nhiờn v a dng sinh hc Vai trũ ca n v bo tn l m nhim vai trũ tr giỳp v k thut, kinh t, cung cp trang thit b cho i ng cỏn b a phng, ng thi tham gia h tr trc tip, giỏm sỏt, ỏnh giỏ kt qu ngi dõn thc hin Trong cụng tỏc giỏo dc tuyờn truyn, nờn trỳ trng cung cp ti liu, tranh nh v mụ hỡnh thc tin, trung vo i tng cn bo tn + Gii phỏp v bo tn ngoi vi: u tiờn xõy dng cỏc mụ hỡnh sinh k v nhõn nuụi, phỏt trin, tiu d ỏn nh v thc hin ti a phng- c bit trung cỏc loi cú giỏ tr kinh t cao, cú s lng ớt, phõn b hp Trong d ỏn, mụ hỡnh sinh k, n v bo tn úng vai trũ c k thut, xõy dng, ch o v cựng thc hin, ỏnh giỏ v tng kt mụ hỡnh + Gii phỏp v bo tn ni vi: cỏn b Kim lõm cn tng cng kt hp vi chớnh quyn a phng tham gia trc tip vo vic tra, kim soỏt khai thỏc cỏc loi c cn lm thng phm, thc phm; ng thi ngn chn trit tỡnh trng khai thỏc g, phỏt nng lm ry lm nh hng ti che ph v tớnh a dng thc vt trờn nỳi ỏ Vic kim soỏt nờn thc hin cựng vi chng trỡnh giỏm sỏt a dng v trung vo cỏc loi cú s lng ớt, cú giỏ tr kinh t cao + Gii phỏp v khoa hc: nghiờn cu v a dng sinh hc, sinh hc, sinh thỏi ca cỏc loi c cn, c bit l cỏc loi cú giỏ tr kinh t, s lng ớt, phõn b hp nờn c u tiờn nhm hon thin thụng tin, hiu bit v chỳng; ng thi tin hnh vic xõy dng bo tn gen iu kin ụng lnh sõu lu tr lõu di s l hng nghiờn cu, bo tn an ton cho tng lai Cỏc nghiờn cu ny cn cú s tham gia ca cỏc n v hng u th gii lnh vc nghiờn cu sinh hc v bo tn Sofhúa bi trung tõm hc liu Footer Page 96 126 http://www.lrc.tnu.edu.vn/ Header Page 97 of 126 89 TI LIU THAM KHO I Ti liu ting Vit Nguyn Th Cy (2006), Bc u nghiờn cu Thõn mm Chõn bng trờn cn khu vc th trn Tam o, Vnh Phỳ, Lun Thc s khoa hc sinh hc, Trng i hc S phm H Ni Nguyn Chớnh (1996), Mt s loi ng vt nhuyn th (Mollusca) cú giỏ tr kinh t bin Vit Nam, Nh xut bn Khoa hc v K thut, H Ni, tr 15-56 inh Phng Dung 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http://www.khoahoc.com.vn/khampha/the-gioi-dong-vat/4173_18-loai-dongvat-khong-xuong-song-o-can-xam-hai-nguy-hiem-nhat-tren-the-gioi.aspx 173 http://www.yhshells.com/goods.php?id=4636 174 Louise R Page, (2006) "Modern insights on gastropod development: Reevaluation of the evolution of a novel body plan", Integrative and Comparative Biology 46 (2): 134143 doi:10.1093/icb/icj018 PMID 21672730 175 Nordsieck H (2010b), Clausiliidae Clausiliidae of Vietnam (Gastropoda, Stylommatophora) Published in Internet: http://www.hnords.e/5356429b13b3201/ index.html S húa bi trung tõm hc liu Footer Page 107 of 126 http://www.lrc.tnu.edu.vn/ Header Page 108 of 126 100 DANH MC CễNG TRèNH CễNG B Khut ng Long, Nguyn Th Liờn, Lng Vn Ho, Vn Lp, 2007 Kho sỏt v ong v ong Bp cy (Hymenoptera) ti VQG Cỳc Phng Bỏo cỏo ti hi ngh khoa hc Quc gia ln th v sinh thỏi v ti nguyờn sinh hc, NXB Nụng nghip, trang 415-419 Masaya Yago, Lng Vn Ho, H Quang Hựng v cng s, 2007 c cn cú vy Bc Vit Nam Phn 2: VQG Cỳc Phng, Ninh Bỡnh Tp hip hi Malacilogical Nht Bn Ting Nht Lng Vn Ho, 2006 Rng nhit i, Phn Chng trỡnh giỏo dc mụi trng trờn VTV2 Lng Vn Ho, 2005 Rng nhit i, Phn Chng trỡnh giỏo dc mụi trng trờn VTV2 S-K, Jemong K-Y, Sung J-M v Lng Vn Ho, 2005 a dng ụng trựng h tho VQG Cỳc Phng, Vit Nam Bỏo cỏo ca i hc Kangwon Ting Anh v ting Hn Trn Vn Móo (ch biờn), Trng Quang Bớch, Vn Lp, Trn Tun Kha, Lng Vn Ho, Lờ Trng t v Lng Vn Hin, 2005 Nm ln Cỳc Phng (Danh sỏch minh ha) Nh xut bn Nụng Nghip H Ni Ting Vit Torben B Larsen, Lng Vn Ho v Vn Lp, 2005 a dng bm ti Vn Quc gia Cỳc Phng, Vit Nam Tp a dng sinh hc, ISSN 1488-8236, Canada Ting Anh Lng Vn Ho (Ch biờn), ng Th ỏp, Trng Quang Bớch v Vn Lp, 2004 Danh lc minh cỏc loi bm Vn Quc gia Cỳc Phng Nh xut bn Nụng nghip, H Ni Ting Vit v Ting Anh S húa bi trung tõm hc liu Footer Page 108 of 126 http://www.lrc.tnu.edu.vn/ ... Bƣởi thuộc Thạch Thành, Thạch Lâm (Thanh Hóa), VQG Cúc Phƣơng Vì vậy, lựa chọn đề tài Nghiên cứu thành phần loài đặc điểm phân bố Thân mềm Chân bụng (Gastropoda) cạn rừng Thạch Thành (Thanh Hóa). .. thành phần đa dạng loài Thân mềm Chân bụng cạn khu rừng Thạch Lâm, Thạch Thành (Thanh Hóa) thuộc Vƣờn Quốc gia Cúc Phƣơng; + Tìm hiểu đặc điểm phân bố chúng sinh cảnh địa hình thuộc khu vực nghiên. .. nghiên cứu; Nội dung 3: Tìm hiểu giá trị thực tiễn Thân mềm Chân bụng cạn Thạch Lâm, Thạch Thành (Thanh Hóa) 2.3.1 Phương pháp nghiên cứu Để thực đề tài Nghiên cứu thành phần loài đặc điểm phân